Kingdom

Shift type

Number of genes

 min max

EUK+1

Eukaryote

 +1

37

–1302

4250

PRO+1

Prokaryote

 +1

50

–1146

1117

EUK−1

Eukaryote

 -1

27

–2849

4170

PRO−1

Prokaryote

 -1

27

–1451

953
Table 2: The four studied frameshift genes populations. Kingdom, shift type, number of genes, minimum i position min and maximum i position max of the studied frameshift gene populations extracted from the RECODE 2 database (release 2010) are given.