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Born in Bloemfontein, South Africa. He graduated in Dentistry from the University of Witwatersrand in 1956. Appointed research fellow at the University of Rochester from 1959 to 1961, where he earned a MS degree. He came to the University of Alberta in 1961 and was a full professor in Dentistry until 1996. He is now an emeritus professor and has been appointed successively as a Visiting Professor at the National University of Singapore, the University of Witwatersrand, University of Otago, the University of Manitoba, and the University of Rochester during the past 14 years.
Human Genetics & Clinical Embryology.
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