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Electron transfer from the cytochrome bf complex to photosystem I is mediated by a small Cu-protein, plastocyanin (PC). Plastocyanin is water soluble and operates in the inner water space of the photosynthetic membrane. Electron transfer from photosystem I to NADP+ requires ferredoxin, a small FeS protein, and ferredoxin-NADP oxidoreductase, a peripheral flavoprotein that operates on the outer surface of the photosynthetic membrane. Ferredoxin and NADP+ are water soluble and are found in the outer aqueous phase.
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