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(Wolf) was born in Berlin, Germany in 1943 and was brought up in the Black Forest region of south-western Germany. He emigrated to Canada at the age of 10 and was schooled in Victoria and Montreal. He graduated from McGill University with B.Sc. and M. Sc. Degrees in Geology, Psychology and Zoology. His first publications dealt with the development of song in cardinal birds. He graduated from the University of Maryland with a Ph.D. in 1974 based on field research concerning the behavior and ecology of Sri Lankan toque macaques. After a brief stint of field research with sloths in Panama, he returned to Sri Lanka and developed an interdisciplinary program of research on toque macaques, hanuman and Purple-faced langurs. He is affiliated with the Smithsonian Conservation Biology Institute in the USA and the Institute of Fundamental Studies in Sri Lanka. He is married and has two children.
Primate behavioral ecology and sociobiology. Primate communication. Environmental conservation Old World primate behavior.
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